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In some fish, capillary blood flows in the opposite direction to the water, causing countercurrent exchange. The muscles on the sides of the pharynx push the oxygen-depleted water out the gill openings. In bony fish, the pumping of oxygen-poor water is aided by a bone that surrounds the gills called the operculum. [6]
The shared trait of breathing via gills in bony fish and cartilaginous fish is a famous example of symplesiomorphy. Bony fish are more closely related to terrestrial vertebrates, which evolved out of a clade of bony fishes that breathe through their skin or lungs, than they are to the sharks, rays, and the other cartilaginous fish. Their kind ...
The spiracle is often located towards the top of the animal allowing breathing even while the animal is mostly buried under sediments. [5] As sharks adapted a faster moving lifestyle some became obligate ram ventilators , breathing exclusively by forcing water through their gills by swimming; among these are requiem sharks and hammerhead sharks ...
Each gill is supported by a cartilaginous or bony gill arch, [1] which helps to maintain the gill's surface area. Bony fish (osteichthyans, mostly teleost ray-finned fish) have four pairs of arches, cartilaginous fish (chondrichthyans) have five to seven pairs, and the more basal jawless fish ("agnathans") have up to seven.
At this point, the free oxygen in the arterial capillaries diffuses into the swim bladder via the gas gland. [50] Unlike bony fish, sharks do not have gas-filled swim bladders for buoyancy. Instead, sharks rely on a large liver filled with oil that contains squalene, and their cartilage, which is about half the normal density of bone. [51]
The gill arches of bony fish typically have no septum, so the gills alone project from the arch, supported by individual gill rays. Some species retain gill rakers. Though all but the most primitive bony fish lack spiracles, the pseudobranch associated with them often remains, being located at the base of the operculum.
Osteichthyes (/ ˌ ɒ s t iː ˈ ɪ k θ iː z / ost-ee-IK-theez), [2] also known as osteichthyans or commonly referred to as the bony fish, is a diverse superclass of vertebrate animals that have endoskeletons primarily composed of bone tissue.
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ in bony fish (but not cartilaginous fish [1]) that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift via swimming, which expends more energy. [2]