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The epidermis is the outermost cell layer of the primary plant body. In some older works the cells of the leaf epidermis have been regarded as specialized parenchyma cells, [1] but the established modern preference has long been to classify the epidermis as dermal tissue, [2] whereas parenchyma is classified as ground tissue. [3]
Anatomy of a eudicot leaf The plant cuticle is a layer of lipid polymers impregnated with waxes that is present on the outer surfaces of the primary organs of all vascular land plants. It is also present in the sporophyte generation of hornworts , and in both sporophyte and gametophyte generations of mosses . [ 2 ]
The epidermis tissue includes several differentiated cell types; epidermal cells, epidermal hair cells , cells in the stomatal complex; guard cells and subsidiary cells. The epidermal cells are the most numerous, largest, and least specialized and form the majority of the epidermis. They are typically more elongated in the leaves of monocots ...
Stoma in a tomato leaf shown via colorized scanning electron microscope image A stoma in horizontal cross section The underside of a leaf. In this species (Tradescantia zebrina) the guard cells of the stomata are green because they contain chlorophyll while the epidermal cells are chlorophyll-free and contain red pigments.
The genetics of stomatal development can be directly studied by imaging of the leaf epidermis using a microscope. Several major control proteins that function in a pathway mediating the development of guard cells and the stomatal pores have been identified. [35] [44]
Parenchyma cells have a variety of functions: In leaves, they form two layers of mesophyll cells immediately beneath the epidermis of the leaf, that are responsible for photosynthesis and the exchange of gases. [2] These layers are called the palisade parenchyma and spongy mesophyll. Palisade parenchyma cells can be either cuboidal or elongated.
The companion cells, connected to the sieve tubes via plasmodesmata, are responsible for loading the phloem with sugars. The bryophytes lack phloem, but moss sporophytes have a simpler tissue with analogous function known as the leptome. This is an electron micrograph of the epidermal cells of a Brassica chinensis leaf. The stomates are also ...
Bulliform cells or motor cells are large, bubble-shaped epidermal cells that occur in groups on the upper surface of the leaves of many monocots. These cells are present on the upper surface of the leaf. They are generally present near the mid-vein portion of the leaf and are large, empty and colourless.