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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
English: xylem (blue) carries water from the roots upwards phloem (orange) carries products of photosynthesis from the place of their origin (source) to organs where they are needed (roots, storage organs, flowers, fruits – sink); note that e.g. the storage organs may be source and leaves may be sink at the beginning of the growing season
The phloem tissue arises from the cell facing outside and consists of sieve tubes and their companion cells. The function of phloem tissue is to distribute food from photosynthetic tissue to other tissues. The two tissues are separated by cambium, a tissue that divides to form xylem or phloem cells.
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Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. There is also a tissue between xylem and phloem, which is the cambium. The xylem typically lies towards the axis with phloem positioned away from the axis . In a stem or root this means that the xylem is closer to the centre of ...
The earliest vascular plants had stems with a central core of vascular tissue. [3] [4] This consisted of a cylindrical strand of xylem, surrounded by a region of phloem. Around the vascular tissue there might have been an endodermis that regulated the flow of water into and out of the vascular system. Such an arrangement is termed a protostele.
Phloem is a nutrient-conducting tissue composed of sieve tubes or sieve cells mixed with parenchyma and fibers. The cortex is the primary tissue of stems and roots. In stems the cortex is between the epidermis layer and the phloem, in roots the inner layer is not phloem but the pericycle .
The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.