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The superior olivary complex (SOC) or superior olive is a collection of brainstem nuclei that is located in pons, functions in multiple aspects of hearing and is an important component of the ascending and descending auditory pathways of the auditory system.
The olivocochlear system is a component of the auditory system involved with the descending control of the cochlea.Its nerve fibres, the olivocochlear bundle (OCB), form part of the vestibulocochlear nerve (VIIIth cranial nerve, also known as the auditory-vestibular nerve), and project from the superior olivary complex in the brainstem to the cochlea.
The olive consists of two parts: The inferior olivary nucleus (or 'complex'), which is a part of the olivo-cerebellar system and is mainly involved in cerebellar motor-learning and function. The superior olivary nucleus, considered part of the pons and part of the auditory system, aiding the perception of sound.
1.2.1 Pons. 1.2.2 Fourth ... Medial superior olive; Lateral superior olive; Medial nucleus of the trapezoid body; ... Cross-section of the midbrain. Tectum. Corpora ...
This marks the end of the pons as the inferior colliculus is directly rostral and marks the caudal midbrain. Middle cerebellar peduncle is located inferior and lateral to the superior cerebellar peduncle, connecting pons to the cerebellum. Likewise, inferior cerebellar peduncle is found connecting the medulla oblongata to the cerebellum.
Diagram of the midbrain, sectioned at the level of the superior colliculus (Central tegmental tract not labeled, but region is visible.) Axial section of the Brainstem (Pons) at the level of the Facial Colliculus (Central tegmental tract not labeled, but region is visible.)
Globular bushy cells project large axons to the contralateral medial nucleus of the trapezoid body (MNTB), in the superior olivary complex where they synapse onto principal cells via a single calyx of Held, and several smaller collaterals synapse ipsilaterally in the posterior (PPO) and dorsolateral periolivary (DLPO) nuclei, lateral superior olive (LSO), and lateral nucleus of the trapezoid ...
Fibers leaving these brainstem nuclei ascending to the inferior colliculus rejoin the lateral lemniscus. In that sense, this is not a 'lemniscus' in the true sense of the word (second order, decussated sensory axons), as there is third (and out of the lateral superior olive, fourth) order information coming out of some of these brainstem nuclei.