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A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells.
Most true fungi have a cell wall consisting largely of chitin and other polysaccharides. [28] True fungi do not have cellulose in their cell walls. [16] In fungi, the cell wall is the outer-most layer, external to the plasma membrane. The fungal cell wall is a matrix of three main components: [16]
The cell wall (3) swells around the septal pore to form a barrel-shaped ring. Perforations in the parenthesome allow cytoplasm to flow between (4) and (5). Dolipore septa are specialized dividing walls between cells (septa) found in almost all species of fungi in the phylum Basidiomycota . [ 1 ]
Typical fungal cell wall structure. Zygomycetes exhibit a special structure of cell wall. Most fungi have chitin as structural polysaccharide, while zygomycetes synthesize chitosan, the deacetylated homopolymer of chitin. Chitin is built of β-1,4 bonded N-acetyl glucosamine. Fungal hyphae grow at the tip.
A clamp connection is a hook-like structure formed by growing hyphal cells of certain fungi. It is a characteristic feature of basidiomycete fungi. It is created to ensure that each cell , or segment of hypha separated by septa (cross walls), receives a set of differing nuclei , which are obtained through mating of hyphae of differing sexual types.
Microfungi or micromycetes are fungi—eukaryotic organisms such as molds, mildews and rusts—which have microscopic spore-producing structures. [1] They exhibit tube tip-growth and have cell walls composed of chitin, a polymer of N-acetylglucosamine.
The fungal cell wall is made of a chitin-glucan complex; while glucans are also found in plants and chitin in the exoskeleton of arthropods, [36] fungi are the only organisms that combine these two structural molecules in their cell wall. Unlike those of plants and oomycetes, fungal cell walls do not contain cellulose. [37] [38]
Chitin-glucan complex (CGC) is a copolymer (polysaccharide) that makes up fungal cell walls, consisting of covalently-bonded chitin and branched 1,3/1,6-ß-D-glucan. CGCs are alkaline-insoluble. Different species of fungi have different structural compositions of chitin and β-glucan making up the CGCs in their cell walls. [1]