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In biology, electrochemical gradients allow cells to control the direction ions move across membranes. In mitochondria and chloroplasts, proton gradients generate a chemiosmotic potential used to synthesize ATP, [1] and the sodium-potassium gradient helps neural synapses quickly transmit information. [citation needed]
Both organelles, the mitochondria and chloroplasts (in photosynthetic organisms), are compartments that are believed to be of endosymbiotic origin. Other compartments such as peroxisomes , lysosomes , the endoplasmic reticulum , the cell nucleus or the Golgi apparatus are not of endosymbiotic origin.
Eukaryotic cells contain organelles including mitochondria, which provide energy for cell functions; chloroplasts, which create sugars by photosynthesis, in plants; and ribosomes, which synthesise proteins. Cells were discovered by Robert Hooke in 1665, who named them after their resemblance to cells inhabited by Christian monks in a monastery.
Mitochondria and plastids contain their own ribosomes; these are more similar to those of bacteria (70S) than those of eukaryotes. [74] Proteins created by mitochondria and chloroplasts use N-formylmethionine as the initiating amino acid, as do proteins created by bacteria but not proteins created by eukaryotic nuclear genes or archaea. [75] [76]
In cell biology, this type of signaling typically occurs between the mitochondria or chloroplast and the nucleus. Signaling molecules from the mitochondria or chloroplast act on the nucleus to affect nuclear gene expression. In this regard, the chloroplast or mitochondria act as a sensor for internal external stimuli which activate a signaling ...
Three general types of extranuclear inheritance exist. Vegetative segregation results from random replication and partitioning of cytoplasmic organelles. It occurs with chloroplasts and mitochondria during mitotic cell divisions and results in daughter cells that contain a random sample of the parent cell's organelles.
In mitochondria, the PMF is almost entirely made up of the electrical component but in chloroplasts the PMF is made up mostly of the pH gradient because the charge of protons H + is neutralized by the movement of Cl − and other anions. In either case, the PMF needs to be greater than about 460 mV (45 kJ/mol) for the ATP synthase to be able to ...
Unlike the IMS of the mitochondria, the IMS of the chloroplast does not seem to have any obvious function. The translocase of the outer membrane and the translocase of the inner membrane mainly assist the translocation of chloroplast precursor proteins [10] Chaperone involvement in the IMS has been proposed but still remains uncertain.