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Psilotum is a genus of fern-like vascular plants.It is one of two genera in the family Psilotaceae commonly known as whisk ferns, the other being Tmesipteris.Plants in these two genera were once thought to be descended from the earliest surviving vascular plants, but more recent phylogenies place them as basal ferns, as a sister group to Ophioglossales.
The scales are arranged in two rows along the flat stems and branches. The stems are broadly triangular in cross section and 5 mm wide. The branches are 1.5 to 2 mm wide. P. complanatum grows 10 to 75 cm long and stems branch in pairs a number of times up their length and are covered with brownish colored hair-like rhizoids.
Higher up it became elliptical in cross-section, corresponding to the two rows of fertile branching units. Within the branches of these units, the strand was more-or-less rectangular. The conducting elements of the xylem, the tracheids , were of the so-called 'P-type' in which the walls were strengthened by ladder-like (scalariform) bars with ...
An interesting case is that of Psilotum, which has a (simple) protostele, and enations devoid of vascular tissue. Some species of Psilotum have a single vascular trace that terminates at the base of the enations. [2] Consequently, Psilotum was long thought to be a "living fossil" closely related to early land plants (rhyniophytes).
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Cross-section of a flax plant stem: 1. Pith 2. Protoxylem 3. Xylem I 4. Phloem I 5. Sclerenchyma 6. Cortex 7. Epidermis. In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. [1]
F bicollateral open Cross section of celery stalk, showing vascular bundles, which includes both phloem and xylem Detail of vascular bundle: closed, collateral vascular bundles of the stem axis of Zea mays Vascular bundle in the leaf of Metasequoia glyptostroboides The vascular bundle of pine leaf showing xylem and phloem
Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. [2]