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Paenungulata (from Latin paene "almost" + ungulātus "having hoofs") is a clade of "sub-ungulates", which groups three extant mammal orders: Proboscidea (including elephants), Sirenia (sea cows, including dugongs and manatees), and Hyracoidea ().
Proboscidea (/ ˌ p r oʊ b ə ˈ s ɪ d i ə /; from Latin proboscis, from Ancient Greek προβοσκίς (proboskís) 'elephant's trunk') is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families.
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Primelephas is a genus of Elephantinae [1] that existed during the Miocene and Pliocene epochs. The name of the genus suggests 'first elephant'. These primitive elephantids are thought to be the common ancestor of Mammuthus, the mammoths, and the closely allied genera Elephas and Loxodonta, the Asian and African elephants, diverging some 4-6 million years ago. [2]
The earliest members of the modern genera of Elephantidae appeared during the latest Miocene–early Pliocene around 5 million years ago. The elephantid genera Elephas (which includes the living Asian elephant) and Mammuthus (mammoths) migrated out of Africa during the late Pliocene, around 3.6 to 3.2 million years ago. [17]
Molar of a modern African elephant for comparison Early gomphotheres had lower jaws with an elongate (longitostrine) mandibular symphysis (the front-most part of the lower jaw) and lower tusks, the primitive condition for members of Elephantimorpha .
The ancestry from L. cyclotis was more closely related to modern West African populations of the forest elephant than to other forest elephant populations, while the mammoth ancestry was basal to the split between woolly and Columbian mammoths, probably from shortly after the split between the ancestors of mammoths and Asian elephants.
Evolutionary and demographic processes shaping geographic patterns of genetic diversity in a keystone species, the African forest elephant (Loxodonta cyclotis). Ecology and Evolution. Stéphanie Bourgeois et al. (2018). Single-nucleotide polymorphism discovery and panel characterization in the African forest elephant. Ecology and Evolution.