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Cyanobacteria use photosynthetic pigments such as various forms of chlorophyll, carotenoids, phycobilins to convert the photonic energy in sunlight to chemical energy. Unlike heterotrophic prokaryotes, cyanobacteria have internal membranes. These are flattened sacs called thylakoids where photosynthesis is performed.
Cyanobacteria is the only prokaryotic group that performs oxygenic photosynthesis. Anoxygenic photosynthetic bacteria use PSI- and PSII-like photosystems, which are pigment protein complexes for capturing light. [5] Both of these photosystems use bacteriochlorophyll. There are multiple hypotheses for how oxygenic photosynthesis evolved.
Photoheterotrophs generate ATP using light, in one of two ways: [6] [7] they use a bacteriochlorophyll-based reaction center, or they use a bacteriorhodopsin.The chlorophyll-based mechanism is similar to that used in photosynthesis, where light excites the molecules in a reaction center and causes a flow of electrons through an electron transport chain (ETS).
In an ecological context, phototrophs are often the food source for neighboring heterotrophic life. In terrestrial environments, plants are the predominant variety, while aquatic environments include a range of phototrophic organisms such as algae (e.g., kelp), other protists (such as euglena), phytoplankton, and bacteria (such as cyanobacteria).
Cyanobacteria such as these carry out photosynthesis. Their emergence foreshadowed the evolution of many photosynthetic plants and oxygenated Earth's atmosphere. Biological carbon fixation, or сarbon assimilation, is the process by which living organisms convert inorganic carbon (particularly carbon dioxide) to organic compounds.
As cyanobacteria are bacteria that use light to fuel their energy-producing photosynthetic machinery they depend on perceiving light in order to optimize their response and to avoid harmful light that could result in the formation of reactive oxygen species and subsequently in their death. [28]
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[12] [27] Cyanobacteria benefit their hosts through providing glycerol and organic phosphates through photosynthesis and supply up to half of their required energy and a majority of their carbon budget. [28] Two groups of sponges with photosynthetic symbionts have been described; these are the "cyanosponges" and "phototrophs".