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Spongy tissue is a type of tissue found both in plants and animals. In plants, it is part of the mesophyll , where it forms a layer next to the palisade cells in the leaf . The spongy mesophyll's function is to allow for the interchange of gases (CO 2 ) that are needed for photosynthesis .
Parenchyma cells have a variety of functions: In leaves, they form two layers of mesophyll cells immediately beneath the epidermis of the leaf, that are responsible for photosynthesis and the exchange of gases. [2] These layers are called the palisade parenchyma and spongy mesophyll. Palisade parenchyma cells can be either cuboidal or elongated.
Palisade cells are located beneath the upper epidermis and cuticle but above the spongy mesophyll cells. Palisade cells contain a high concentration of chloroplasts , particularly in the upper portion of the cell, making them the primary site of photosynthesis in the leaves of plants that contain them.
The pores or stomata of the epidermis open into substomatal chambers, which are connected to the intercellular air spaces between the spongy and palisade mesophyll cell, so that oxygen, carbon dioxide and water vapor can diffuse into and out of the leaf and access the mesophyll cells during respiration, photosynthesis and transpiration.
Aerenchyma in stem cross section of a typical wetland plant. Aerenchyma or aeriferous parenchyma [1] or lacunae, is a modification of the parenchyma to form a spongy tissue that creates spaces or air channels in the leaves, stems and roots of some plants, which allows exchange of gases between the shoot and the root. [2]
The primary function of kranz anatomy is to provide a site in which CO 2 can be concentrated around RuBisCO, thereby avoiding photorespiration. Mesophyll and bundle sheath cells are connected through numerous cytoplasmic sleeves called plasmodesmata whose permeability at leaf level is called bundle sheath conductance.
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As a result, chloroplasts in C 4 mesophyll cells and bundle sheath cells are specialized for each stage of photosynthesis. In mesophyll cells, chloroplasts are specialized for the light reactions, so they lack RuBisCO, and have normal grana and thylakoids, [140] which they use to make ATP and NADPH