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Please do so irrespective of any rules associated with the variety of English in use. The serial comma (for example the comma before and in "ham, chips, and eggs") is optional; be sensitive to possible ambiguity arising from thoughtless use or thoughtless avoidance, and be consistent within a given article. Avoid comma splices.
In the actual title of a published work, like the "Luke Skywalker: A pilot in the Rebel Alliance and apprentice Jedi" example given above, capitalization would follow the colon because there we are dealing with a subtitle, and we could not replace the colon with a comma. In short, we already capitalize after a colon when something is a subtitle ...
It is a simple of matter of fact that people do capitalize titles when they stand in the place of names, just as other people do not. People do capitalize every instance of a title, just as other people put nearly everyone in lower case. This a question of style. As such, it should reflect best practices and actual usage.
Do not ordinarily capitalize the definite article after the first word of a sentence; [a] however, some idiomatic expressions, including the titles of artistic and academic works, should be quoted exactly, according to common usage. Correct (generic): an article about the United Kingdom Incorrect: an article about The United Kingdom (a redirect)
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The initial lowercase is often there to indicate reading as a letter, as opposed to as a part of a work, as in eBay, iPod. In the case of WatchOS, no initial cap is needed, and in fact some book sources do capitalize it, so we're choosing from among existing styles if we do, too (for title and sentence initial, or maybe always). So that leaves ...
As SchreiberBike said, we do so because that's the accepted style; first words and last words are more significant than middle words in titles, and are capitalized even whey they are short prepositions, conjunctions, or other words that wouldn't be capitalized in the middle of a title. -- JHunterJ 18:53, 7 April 2014 (UTC)
The simplest de-capitalization rule is to capitalize if, and only if, the title is directly used as a title in front of a name, so "President Nixon" but everywhere else "president". Such a rule could actually be followed.