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Some prokaryotes are pathogenic, causing disease and even death in plants and animals. [5] Marine prokaryotes are responsible for significant levels of the photosynthesis that occurs in the ocean, as well as significant cycling of carbon and other nutrients. [6] Prokaryotes live throughout the biosphere.
They can occur as planktonic cells or form phototrophic biofilms. They are found inside stones and shells (in endolithic ecosystems). [95] A few are endosymbionts in lichens, plants, various protists, or sponges and provide energy for the host. Some live in the fur of sloths, providing a form of camouflage. [96]
Cyanobacteria is the only prokaryotic group that performs oxygenic photosynthesis. Anoxygenic photosynthetic bacteria use PSI- and PSII-like photosystems, which are pigment protein complexes for capturing light. [5] Both of these photosystems use bacteriochlorophyll. There are multiple hypotheses for how oxygenic photosynthesis evolved.
Some dinoflagellates are known to be photosynthetic, but a large fraction of these are in fact mixotrophic, combining photosynthesis with ingestion of prey (phagotrophy). [76] Some species are endosymbionts of marine animals and other protists, and play an important part in the biology of coral reefs .
Some archaeans are methanogens, living in anoxic environments and releasing methane. [2] Many archaea grow as plankton in the oceans. Symbiotic prokaryotes live in or on the bodies of other organisms, including humans. Prokaryotes have high populations in the soil, in the sea, and in undersea sediments.
Heliobacteria are a unique subset of prokaryotic bacteria that process light for energy. Distinguishable from other phototrophic bacteria, they utilize a unique photosynthetic pigment, bacteriochlorophyll g and are the only known Gram-positive phototroph. [ 2 ]
Cyanobacteria, the only prokaryotes performing oxygenic photosynthesis and the only prokaryotes that contain two types of photosystems (type I (RCI), also known as Fe-S type, and type II (RCII), also known as quinone type). The seven remaining prokaryotes have anoxygenic photosynthesis and use versions of either type I or type II.
Photoheterotrophs generate ATP using light, in one of two ways: [6] [7] they use a bacteriochlorophyll-based reaction center, or they use a bacteriorhodopsin.The chlorophyll-based mechanism is similar to that used in photosynthesis, where light excites the molecules in a reaction center and causes a flow of electrons through an electron transport chain (ETS).
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