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The first of the cells (the generative cell) is small and is formed inside the second larger cell (the tube cell). The members of each part of the microspores separate from each other. A double-layered wall then develops around each microspore. These steps occur in sequence and when complete, the microspores have become pollen grains. [3]
The pollen grain surface is covered with waxes and proteins, which are held in place by structures called sculpture elements on the surface of the grain. The outer pollen wall, which prevents the pollen grain from shrinking and crushing the genetic material during desiccation, [citation needed] is composed of two layers. These two layers are ...
These cells are surrounded by a wall of sterile cells called the tapetum, which supplies food to the cell and eventually becomes the cell wall for the pollen grain. These sets of sporogenous cells eventually develop into diploid microspore mother cells. These microspore mother cells, also called microsporocytes, then undergo meiosis and become ...
SEM image of pollen grains. Sporopollenin is a biological polymer found as a major component of the tough outer (exine) walls of plant spores and pollen grains. It is chemically very stable (one of the most inert among biopolymers) [1] and is usually well preserved in soils and sediments.
However, with the development of the electron microscope, many of the questions were answered. Most notably, the observations made by the group of W. Jensen showed that the male gametes did not have any cell walls and that the plasma membrane of the gametes is close to the plasma membrane of the cell that surrounds them inside the pollen grain. [6]
The cells of the primary parietal layer divide by successive periclinal and anticlinal divisions to form concentric layers of pollen sac wall. [citation needed] The wall layers from periphery to center consist of: [clarification needed] A single layer of epidermis, which becomes stretched and shrivels off at maturity; A single layer of ...
The tapetum is a specialised layer of nutritive cells found within the anther of flowering plants, located between the sporogenous tissue and the anther wall. Tapetum is important for the nutrition and development of pollen grains and a source of precursors for the pollen coat. [1]
Spore mother cells in the microsporangia divide by meiosis to form haploid microspores that develop further by two mitotic divisions into immature male gametophytes (pollen grains). The four resulting cells consist of a large tube cell that forms the pollen tube, a generative cell that will produce two sperm by mitosis, and two prothallial ...