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Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and Mollisonia present by this time. [2] The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates.
The subphylum Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes), and all true insects.
The jumping spider Phidippus audax.The basal parts of the chelicerae are the two iridescent green mouthparts. The chelicerae (/ k ə ˈ l ɪ s ər iː /) are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders.
Paired maxillae cut food and manipulate it during mastication. Maxillae can have hairs and "teeth" along their inner margins. At the outer margin, the galea is a cupped or scoop-like structure, which sits over the outer edge of the labium. They also have palps, which are used to sense the characteristics of potential foods.
However, a 2022 analysis recovered the monophyly of Merostomata, as opposed to a monophyletic Arachnida, with Xiphosura as the only modern representative in a derived position, indicating the convergence of several characteristics that supposedly united the arachnids.
The Crustacea were generally considered the closest relatives of the Uniramia, and sometimes these were united as Mandibulata.However, the competing hypothesis — that Crustacea and Hexapoda form a monophyletic group, the Pancrustacea, to which the Myriapoda are the closest relatives — has support from molecular and fossil evidence.
The species formerly placed in Haplopelma are medium to large spiders; for example, Cyriopagopus schmidti females have a total body length, including chelicerae, up to 85 mm (3.3 in), with the longest leg, the first, being about 70 mm (2.8 in) long.
Schizomida, also known as sprickets [1] or short-tailed whip-scorpions, [2] is an order of arachnids, generally less than 5 millimetres (0.20 in) in length.The order is not yet widely studied.