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The authors favoured relationships shown by more slowly evolving genes, which demonstrated the monophyly of Chelicerata, Euchelicerata and Arachnida, as well as of some clades within the arachnids. The diagram below summarizes their conclusions, based largely on the 200 most slowly evolving genes; dashed lines represent uncertain placements.
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The group's origins lie within an arachnid sub-group defined by the presence of book lungs (the tetrapulmonates); [1] [2] the arachnids as a whole evolved from aquatic chelicerate ancestors. More than 45,000 extant species have been described, organised taxonomically in 3,958 genera and 114 families. [3] There may be more than 120,000 species. [3]
The cephalothorax, also called prosoma, is composed of two primary surfaces: a dorsal carapace and a ventral sternum.Most external appendages on the spider are attached to the cephalothorax, including the eyes, chelicerae and other mouthparts, pedipalps and legs.
The idea of a tree of life arose from ancient notions of a ladder-like progression from lower into higher forms of life (such as in the Great Chain of Being).Early representations of "branching" phylogenetic trees include a "paleontological chart" showing the geological relationships among plants and animals in the book Elementary Geology, by Edward Hitchcock (first edition: 1840).
Close-up of the chelicerae of a harvestman. The chelicerae consist of three segments, with the distal two forming a pincer [3] equipped with a row of teeth. Based on developmental gene expression data, the proximal segment is thought to be homologous to the femur and patella of the legs, the middle segment to the tibia through the tarsus of the legs, and the distal segment to the apotele (the ...