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The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
The three-domain system presumes that eukaryotes are more closely related to archaea than to Bacteria and are sister group to Archaea, thus, it treats them as separate domain. [29] As more new archaea were discovered in the early 2000s, this distinction became doubtful as eukaryotes became deeply nested within Archaea.
The last universal common ancestor (LUCA) is the hypothesized common ancestral cell from which the three domains of life, the Bacteria, the Archaea, and the Eukarya originated. The cell had a lipid bilayer ; it possessed the genetic code and ribosomes which translated from DNA or RNA to proteins .
According to the domain system, the tree of life consists of either three domains, Archaea, Bacteria, and Eukarya, [1] or two domains, Archaea and Bacteria, with Eukarya included in Archaea. [3] [4] In the three-domain model, the first two are prokaryotes, single-celled microorganisms without a membrane-bound nucleus.
The eukaryotes seemingly emerged within the Asgard archaea, and are closely related to the Heimdallarchaeia. [5] This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea.
According to genetic data, although eukaryote groups such as plants, fungi, and animals may look different, they are more closely related to each other than they are to either the Eubacteria or Archaea. It was also found that the eukaryotes are more closely related to the Archaea than they are to the Eubacteria.
Phylogenetic tree showing the relationship between the archaea and other forms of life. Eukaryotes are colored red, archaea green and bacteria blue. Adapted from Ciccarelli et al. [44] Woese argued that the bacteria, archaea, and eukaryotes represent separate lines of descent that diverged early on from an ancestral colony of organisms.
For example, Japanese scientists reported in 1990 their study on the elongation factors Tu(EF-Tu) and G(EF-G) from various organisms that showed that eukaryotes are most closely related to archaea (methanogen and halobacteria), and not eocytes. [15] Other studies also supported the eukaroyte-archaea relationship and rejected the eocyte hypotheses.