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There are two developmental stages beyond the development of the Casparian strip. In the second stage suberin (or endoderm [9]) coats the entire wall on the inside of the cell. As a result, the Casparian strip is separated from the cytoplasm and the connection between the two ceases to be evident.
Suberin is highly hydrophobic and a somewhat 'rubbery' material. In roots, suberin is deposited in the radial and transverse/tangential cell walls of the endodermal cells. This structure, known as the Casparian strip or Casparian band, functions to prevent water and nutrients taken up by the root from entering the stele through the apoplast.
Stage 2 includes the deposition of suberin and other hydrophobic polymers and cell membranes of individual exodermal cells. It also serves in forming the connection between the plasmodesma and the Casparian strip. Stage 3 includes the addition of cellulose and lignin with occasional deposition of suberin into the cell walls to strengthen them. [1]
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If the endodermis is likened to a brick cylinder (e.g. a smokestack), with the bricks representing individual cells, the Casparian strips are analogous to the mortar between the bricks. In older endodermal cells, suberin may be more extensively deposited on all cell wall surfaces and the cells can become lignified, forming a complete waterproof ...
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Casparian strip A continuous band of suberin in the radial primary cell walls of the endodermis in vascular plant stems and roots that forms a permeability barrier to the passive diffusion of external water and solutes into the vascular tissue. cassideous