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The idea of a tree of life arose from ancient notions of a ladder-like progression from lower into higher forms of life (such as in the Great Chain of Being).Early representations of "branching" phylogenetic trees include a "paleontological chart" showing the geological relationships among plants and animals in the book Elementary Geology, by Edward Hitchcock (first edition: 1840).
The science that tries to reconstruct phylogenetic trees and thus discover clades is called phylogenetics or cladistics, the latter term coined by Ernst Mayr (1965), derived from "clade". The results of phylogenetic/cladistic analyses are tree-shaped diagrams called cladograms; they, and all their branches, are phylogenetic hypotheses. [12]
If a phylogenetic tree is reconstructed from DNA sequence data of a particular gene, a hard polytomy arises when three or more sampled genes trace their ancestry to a single gene in an ancestral organism. In contrast, a soft polytomy stems from branches on gene trees of finite temporal duration but for which no substitutions have occurred. [7]
Algorithms for cladograms or other types of phylogenetic trees include least squares, neighbor-joining, parsimony, maximum likelihood, and Bayesian inference. Biologists sometimes use the term parsimony for a specific kind of cladogram generation algorithm and sometimes as an umbrella term for all phylogenetic algorithms. [15]
The results are a phylogenetic tree—a diagram setting the hypothetical relationships between organisms and their evolutionary history. [4] The tips of a phylogenetic tree can be living taxa or fossils, which represent the present time or "end" of an evolutionary lineage, respectively. A phylogenetic diagram can be rooted or unrooted.
Branches down to the divergence to the next significant (e.g. extant) sister are considered stem-groupings of the clade, but in principle each level stands on its own, to be assigned a unique name. For a fully bifurcated tree, adding a group to a tree also adds an additional (named) clade, and a new level on that branch.
[15] Darwin's tree is not a tree of life, but rather a small portion created to show the principle of evolution. Because it shows relationships (phylogeny) and time (generations), it is a timetree. In contrast, Ernst Haeckel illustrated a phylogenetic tree (branching only) in 1866, not scaled to time, and of real species and higher taxa. In his ...
A simple cladogram showing the evolutionary relationships between four species: A, B, C, and D. Here, Species A is the outgroup, and Species B, C, and D form the ingroup. In cladistics or phylogenetics, an outgroup [1] is a more distantly related group of organisms that serves as a reference group when determining the evolutionary relationships of the ingroup, the set of organisms under study ...