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The Lotka–Volterra system of equations is an example of a Kolmogorov population model (not to be confused with the better known Kolmogorov equations), [2] [3] [4] which is a more general framework that can model the dynamics of ecological systems with predator–prey interactions, competition, disease, and mutualism.
Examples include predator-prey competition and host-parasite co-evolution, as well as mutualism. Evolutionary game models have been created for pairwise and multi-species coevolutionary systems. [58] The general dynamic differs between competitive systems and mutualistic systems.
Predators receive a reproductive payoff, e, for consuming prey, and die at rate u. Making predation pressure a function of the ratio of prey to predators contrasts with the prey-dependent Lotka–Volterra equations, where the per capita effect of predators on the prey population is simply a function of the magnitude of the prey population g(N).
The selfish herd theory may also be applied to the group escape of prey in which the safest position, relative to predation risk, is not the central position, but rather the front of the herd. [2] The theory may be useful in explaining the escape strategy chosen by a herd leader. [ 2 ]
Smale [3] showed that Lotka–Volterra systems that meet the above conditions and have five or more species (N ≥ 5) can exhibit any asymptotic behavior, including a fixed point, a limit cycle, an n-torus, or attractors. Hirsch [4] [5] [6] proved that all of the dynamics of the attractor occur on a manifold of dimension N−1.
EcoSim is an individual-based predator-prey ecosystem simulation in which agents can evolve. It has been designed to investigate several broad ecological questions, as well as long-term evolutionary patterns and processes such as speciation and macroevolution.
Predator–prey isoclines before and after pesticide application. Pest abundance has increased. Now, to account for the difference in the population dynamics of the predator and prey that occurs with the addition of pesticides, variable q is added to represent the per capita rate at which both species are killed by the pesticide.
A trophic function was first introduced in the differential equations of the Kolmogorov predator–prey model. It generalizes the linear case of predator–prey interaction firstly described by Volterra and Lotka in the Lotka–Volterra equation. A trophic function represents the consumption of prey assuming a given number of predators.