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Variations are frequent and multiple. In a typical Basidiomycota lifecycle the long lasting dikaryons periodically (seasonally or occasionally) produce basidia, the specialized usually club-shaped end cells, in which a pair of compatible nuclei fuse to form a diploid cell.
Basidiospores typically each contain one haploid nucleus that is the product of meiosis, and they are produced by specialized fungal cells called basidia. Typically, four basidiospores develop on appendages from each basidium, of which two are of one strain and the other two of its opposite strain. In gills under a cap of one common species ...
The phylum Basidiomycota can be divided into three major lineages: mushrooms, rusts and smuts. Fusion of haploid nuclei occurs in the basidia, club-shaped end cells. Shortly after formation of the diploid cell, meiosis occurs and the resulting four haploid nuclei migrate into four, usually external cells called basidiospores.
Most basidiomycota have single celled basidia (holobasidia), but some have ones with many cells (a phragmobasidia).For instance, rust fungi in the order Puccinales have phragmobasidia with four cells that are separated by walls along their cross section.
In the Basidiomycota this is the dominant phase, with most Basidiomycota monokaryons weakly growing and short-lived. Dikaryons shown in a Basidiomycete mitosis cycle The formation of a dikaryon is a plesiomorphic character for the subkingdom Dikarya , which consists of the Basidiomycota and the Ascomycota .
After formation of the ascus-initial cell, the A and a nuclei fuse with each other to form a diploid nucleus (see Figure). This nucleus is the only diploid nucleus in the entire life cycle of N. crassa. The diploid nucleus has 14 chromosomes formed from the two fused haploid nuclei that had 7 chromosomes each.
Coprinopsis cinerea is an ideal model for studying meiosis because meiosis progresses synchronously in about 10 million cells within each mushroom cap. [14] Meiosis is a specialized cell division process, occurring in diploid cells, in which a single round of DNA replication occurs, and is followed by two divisions to produce four haploid ...
All basidiocarps serve as the structure on which the hymenium is produced. Basidia are found on the surface of the hymenium, and the basidia ultimately produce spores. In its simplest form, a basidiocarp consists of an undifferentiated fruiting structure with a hymenium on the surface; such a structure is characteristic of many simple jelly and club fungi.