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In evolutionary biology, punctuated equilibrium (also called punctuated equilibria) is a theory that proposes that once a species appears in the fossil record, the population will become stable, showing little evolutionary change for most of its geological history. [1] This state of little or no morphological change is called stasis.
The color codes the age of the species. The Bak–Sneppen model is a simple model of co-evolution between interacting species. It was developed to show how self-organized criticality may explain key features of the fossil record, such as the distribution of sizes of extinction events and the phenomenon of punctuated equilibrium.
Eldredge and Stephen Jay Gould proposed punctuated equilibria in 1972. Punctuated equilibrium is a refinement to evolutionary theory. It describes patterns of descent taking place in "fits and starts" separated by long periods of stability. Eldredge went on to develop a hierarchical vision of evolutionary and ecological systems. Around this ...
Punctuated gradualism is considered to be a variation of these models, lying somewhere in between the phyletic gradualism model and the punctuated equilibrium model. It states that speciation is not needed for a lineage to rapidly evolve from one equilibrium to another but may show rapid transitions between long-stable states.
Gould argued that punctuated equilibrium challenges the gradual change expected by extrapolationists. In the case of Hominid evolution, there is the evolutionary trend of marked increase in brain size. To Gould, this trend was the result of species sorting, in which species with relatively larger brains were more likely to appear, or to survive.
According to Gould, punctuated equilibrium revised a key pillar "in the central logic of Darwinian theory." [17] Some evolutionary biologists have argued that while punctuated equilibrium was "of great interest to biology generally," [42] it merely modified neo-Darwinism in a manner that was fully compatible with what had been known before. [43]
However, the model of punctuated equilibrium is more widely accepted, with the proposal that most species remain in stasis, a state of very little evolutionary change, for a large proportion of the species' lifespan. [15] This would result in increased cases of speciation through cladogenesis and true extinction, with fewer cases of ...
In an evolutionarily stable state, a population's genetic composition is restored by selection after a disturbance, if the disturbance is not too large. An evolutionarily stable state is a dynamic property of a population that returns to using a strategy, or mix of strategies, if it is perturbed from that initial state.