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An endosymbiont or endobiont [1] is an organism that lives within the body or cells of another organism. Typically the two organisms are in a mutualistic relationship. Examples are nitrogen-fixing bacteria (called rhizobia ), which live in the root nodules of legumes , single-cell algae inside reef-building corals , and bacterial endosymbionts ...
As an endosymbiont evolves into an organelle, most of its genes are transferred to the host cell genome. [24] The host cell and organelle therefore need to develop a transport mechanism that enables the return of the protein products needed by the organelle but now manufactured by the cell.
A symbiosome is formed as a result of a complex and coordinated interaction between the symbiont host and the endosymbiont. [5] At the point of entry into a symbiont host cell, part of the cell's membrane envelops the endosymbiont and breaks off into the cytoplasm as a discrete unit, an organelle-like vacuole called the symbiosome.
As the endosymbiont adapts to the host's lifestyle, the endosymbiont changes dramatically. There is a drastic reduction in its genome size, as many genes are lost during the process of metabolism , and DNA repair and recombination, while important genes participating in the DNA-to-RNA transcription , protein translation and DNA/RNA replication ...
Fungal-bacterial endosymbiosis encompasses the mutualistic relationship between a fungus and intracellular bacteria species residing within the fungus. Many examples of endosymbiotic relationships between bacteria and plants, algae and insects exist and have been well characterized, however fungal-bacteria endosymbiosis has been less well described.
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The hydrogen hypothesis is a model proposed by William F. Martin and Miklós Müller in 1998 that describes a possible way in which the mitochondrion arose as an endosymbiont within a prokaryotic host in the archaea, giving rise to a symbiotic association of two cells from which the first eukaryotic cell could have arisen (symbiogenesis).
The Tsetse fly's most prominent and vital endosymbiont is the bacterium, Wigglesworthia glossinidia, These bacteria are housed in the fly's bacteriocytes and produce B vitamins (B1, B6, and B9). The Tsetse fly itself lacks the ability to obtain these nutrients because of its hematophagous diet.