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Warmer ocean water holds less oxygen and is more buoyant than cooler water. This leads to reduced mixing of oxygenated water near the surface with deeper water, which naturally contains less oxygen. Warmer water also raises oxygen demand from living organisms; as a result, less oxygen is available for marine life. [14]
Upwelling of nutrient-rich water leads to high productivity and labile organic matter, that is respired by heterotrophs as it sinks down the water column. High respiration rates deplete the oxygen in the water column to concentrations of 2 mg/L or less forming the OMZ. [2] OMZs are expanding, with increasing ocean deoxygenation.
An aquatic system lacking dissolved oxygen (0% saturation) is termed anaerobic, reducing, or anoxic. In water, oxygen levels are approximately 7 ppm or 0.0007% in good quality water, but fluctuate. [5] Many organisms require hypoxic conditions. Oxygen is poisonous to anaerobic bacteria for example. [3]
In OMZs oxygen concentration drops to levels <10 nM at the base of the oxycline and can remain anoxic for over 700 m depth. [7] This lack of oxygen can be reinforced or increased due to physical processes changing oxygen supply such as eddy-driven advection, [7] sluggish ventilation, [8] increases in ocean stratification, and increases in ocean temperature which reduces oxygen solubility.
Oxygen diffuses into water from air and therefore the top layer of water in contact with air contains more oxygen. This is true only in stagnant water; in running water all layers are mixed together and oxygen levels are the same throughout the water column. One environment where ASR often takes place is tidepools, particularly at night. [34]
Fountains aerate water by spraying it into the air. Water aeration is the process of increasing or maintaining the oxygen saturation of water in both natural and artificial environments. Aeration techniques are commonly used in pond, lake, and reservoir management to address low oxygen levels or algal blooms.
In some fish, capillary blood flows in the opposite direction to the water, causing countercurrent exchange. The muscles on the sides of the pharynx push the oxygen-depleted water out the gill openings. In bony fish, the pumping of oxygen-poor water is aided by a bone that surrounds the gills called the operculum. [6]
During summer stagnation in stratified, eutrophic lakes, an oxygen deficit develops in the deep water.Increases in nutrient inputs raise the trophic level of these lakes. . The nutrients promote surface algae growth and consequently increases oxygen consumption in the deep zones to match increased amounts of oxidative decomposition of increased amounts of falling organic waste from the surface zon