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The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish.
Gnathostomata is traditionally a infraphylum, broken into three top-level groupings: Chondrichthyes, or the cartilaginous fish; Placodermi, an extinct grade of armored fish; and Teleostomi, which includes the familiar classes of bony fish, birds, mammals, reptiles, and amphibians. Some classification systems have used the term Amphirhina.
Placoderms were among the first jawed fish (their jaws likely evolved from the first pair of gill arches), as well as the first vertebrates to have true teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homologous precursor to hindlimbs in tetrapods.
The upper jaw, or maxilla [12][13] is a fusion of two bones along the palatal fissure that form the upper jaw. This is similar to the mandible (lower jaw), which is also a fusion of two halves at the mandibular symphysis. In bony fish, the maxilla is called the "upper maxilla," with the mandible being the "lower maxilla".
Silurian (443–419 Ma): Many evolutionary milestones occurred during this period, including the appearance of armoured jawless fish, jawed fish, spiny sharks and ray-finned fish. While it is traditional to refer to the Devonian as the age of fishes, recent findings have shown the Silurian was also a period of considerable diversification.
Cladistically included but traditionally excluded taxa. Gnathostomata (jawed vertebrates) Agnatha (/ ˈæɡnəθə, æɡˈneɪθə /; [3] from Ancient Greek ἀ- (a-) 'without' and γνάθος (gnáthos) 'jaws') is an infraphylum [4] of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and ...
Chondrichthyes (/ kɒnˈdrɪkθiiːz /; from Ancient Greek χόνδρος (khóndros) 'cartilage' and ἰχθύς (ikhthús) 'fish') is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have ...
Acanthodii or acanthodians is an extinct class of gnathostomes (jawed fishes). They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans (bony fish) and ...