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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
Since the valuable fibres are located in the phloem, they must often be separated from the woody core, the xylem, and sometimes also from the epidermis. The process for this is retting , and can be performed by micro-organisms either on land (nowadays the most important) or in water, or by chemicals (for instance high pH and chelating agents ...
The phloem is the living portion of the vascular system of a plant, and serves to move sugars and photosynthate from source cells to sink cells. Phloem tissue is made of sieve elements and companion cells, and is surrounded by parenchyma cells. The sieve element cells work as the main player in transport of phloem sap.
Phloem is an equally important plant tissue as it also is part of the 'plumbing system' of a plant. Primarily, phloem carries dissolved food substances throughout the plant. This conduction system is composed of sieve-tube member and companion cells, that are without secondary walls. The parent cells of the vascular cambium produce both xylem ...
In old stems the epidermal layer, cortex, and primary phloem become separated from the inner tissues by thicker formations of cork. Due to the thickening cork layer these cells die because they do not receive water and nutrients. This dead layer is the rough corky bark that forms around tree trunks and other stems.
By studying the phloem of the leaves in vivo through laser microscopy and the usage of fluorescent markers (placed in both companion cells and sieve elements), the network of companion cells with the compact sieve tubes was highlighted. The markers for sieve elements and companion cells was used to study the network and organization of phloem ...
When the water pressure within the xylem reaches extreme levels due to low water input from the roots (if, for example, the soil is dry), then the gases come out of solution and form a bubble – an embolism forms, which will spread quickly to other adjacent cells, unless bordered pits are present (these have a plug-like structure called a ...
The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.