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Species which are particularly characteristic of ancient woodland sites are called ancient woodland indicator species, such as bluebells, ramsons, wood anemone, yellow archangel and primrose for example, representing a type of ecological indicator. [8] Anemonoides nemorosa, the wood anemone
Very rare species are not considered axiophytes; for a species to be a useful indicator of quality habitat it must be relatively frequent in those habitats, but scarce elsewhere. A typical example would be dog's mercury (Mercurialis perennis), a plant slow to colonise new sites, but common in ancient woodland and old hedgerows.
It grows in deciduous woodlands with moist soils, preferring slightly acidic conditions. In the British Isles, colonies are frequently associated with bluebells (Hyacinthoides non-scripta), especially in ancient woodland. It is considered to be an ancient woodland indicator species. [15]
A bluebell wood is a woodland that in springtime has a carpet of flowering bluebells (Hyacinthoides non-scripta) underneath a newly forming leaf canopy. The thicker the summer canopy, the more the competitive ground-cover is suppressed, encouraging a dense carpet of bluebells, whose leaves mature and die down by early summer.
The plant is an ancient woodland indicator, as the ants rarely carry the seeds more than a few yards, seldom crossing a field to go to a new woodland. The Latin specific epithet pratense means "of meadows". [2] Melampyrum pratense is a food plant of the caterpillars of the heath fritillary (Melitaea athalia), a butterfly. [3]
Examples of plant formation include forests, woodlands, and grasslands. Because it may not be possible to determine whether a particular plant was an oak, eucalyptus, or other species, plant formations in paleosols make it possible to identify an ancient woodland ecosystem from an ancient grassland ecosystem. [2]
Some epiphytic lichen species may be used as "ancient woodland indicators"; they can used to quantitatively assess the degree to which a forest has had a long history of canopy continuity. [11] The presence of these species is a reliable indicator that the forest has existed back to early medieval times, without being clear-cut and regrown.
[4] [5] [6] It is a predominantly lowland species that inhabits moist, base-rich, shady habitats including; broad-leaf woodlands, hedgerows, limestone pavements, pastures, and riverbanks. [7] It has been used as an indicator of ancient woodland in East Anglia. [8]