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The gymnosperms (/ ˈ dʒ ɪ m n ə ˌ s p ɜːr m z,-n oʊ-/ ⓘ nə-spurmz, -noh-; lit. ' revealed seeds ') are a group of woody, perennial seed-producing plants, typically lacking the protective outer covering which surrounds the seeds in flowering plants, that include conifers, cycads, Ginkgo, and gnetophytes, forming the clade Gymnospermae [2] The term gymnosperm comes from the ...
Numerous fossil gymnosperm clades once existed that are morphologically at least as distinctive as the four living gymnosperm groups, such as Bennettitales, Caytonia and the glossopterids. When these gymnosperm fossils are considered, the question of gnetophyte relationships to other seed plants becomes even more complicated.
Conifers are the largest and economically most important component group of gymnosperms, but nevertheless they comprise only one of the four groups. The division Pinophyta consists of just one class, Pinopsida, which includes both living and fossil taxa. Subdivision of the living conifers into two or more orders has been proposed from time to time.
Proponents of this theory point out that the gymnosperms have two very similar copies of the gene LFY, while angiosperms just have one. Molecular clock analysis has shown that the other LFY paralog was lost in angiosperms around the same time as flower fossils become abundant, suggesting that this event might have led to floral evolution. [135]
The gymnosperms consist of five orders of seed plants: Cupressales, Cycadales, Ginkgoales, Gnetales and Pinales. [a] They developed more than 350 million years ago, long before flowering plants, according to the fossil record. The name comes from the Greek for "naked seed"; the egg cells are not protected by ovaries, as in flowering plants. [4]
The fossil history of flowering plants records the development of flowers and other distinctive structures of the angiosperms, now the dominant group of plants on land.The history is controversial as flowering plants appear in great diversity in the Cretaceous, with scanty and debatable records before that, creating a puzzle for evolutionary biologists that Charles Darwin named an "abominable ...
Unambiguous fossils of cycads are known from the Early-Middle Permian onwards. [26] Cycads were generally uncommon during the Permian. [27] The two living cycad families are thought to have split from each other sometime between the Jurassic [17] and Carboniferous. [28] Cycads are thought to have reached their apex of diversity during the ...
However, angiosperms appear suddenly and in great diversity in the fossil record in the Early Cretaceous (~130 mya). [48] [49] Claimed records of flowering plants prior to this are not widely accepted. [50] Molecular evidence suggests that the ancestors of angiosperms diverged from the gymnosperms during the late Devonian, about 365 million ...