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Psilotum is a genus of fern-like vascular plants.It is one of two genera in the family Psilotaceae commonly known as whisk ferns, the other being Tmesipteris.Plants in these two genera were once thought to be descended from the earliest surviving vascular plants, but more recent phylogenies place them as basal ferns, as a sister group to Ophioglossales.
A central strand of xylem occupied up to a third of the diameter of the stem. In the lower parts of main stems it was circular, enlarging before it divided prior to dichotomous branching. Higher up it became elliptical in cross-section, corresponding to the two rows of fertile branching units.
An interesting case is that of Psilotum, which has a (simple) protostele, and enations devoid of vascular tissue. Some species of Psilotum have a single vascular trace that terminates at the base of the enations. [2] Consequently, Psilotum was long thought to be a "living fossil" closely related to early land plants (rhyniophytes).
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Parenchyma cells have thin primary walls and usually remain alive after they become mature. Parenchyma forms the "filler" tissue in the soft parts of plants, and is usually present in cortex, pericycle, pith, and medullary rays in primary stem and root. Collenchyma cells have thin primary walls with some areas of secondary thickening ...
It forms a protective covering on the leaf vein and consists of one or more cell layers, usually parenchyma. Loosely-arranged mesophyll cells lie between the bundle sheath and the leaf surface. The Calvin cycle is confined to the chloroplasts of these bundle sheath cells in C 4 plants. C 2 plants also use a variation of this structure. [1]
Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. [2]
Psilotum has been interpreted as producing sporangia (fused in a synangium) on the terminus of a stem. Equisetum always produce strobili, but the structures bearing sporangia (sporangiophores) have been interpreted as modified stems. The sporangia, despite being recurved are interpreted as terminal.