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RuBisCO is important biologically because it catalyzes the primary chemical reaction by which inorganic carbon enters the biosphere.While many autotrophic bacteria and archaea fix carbon via the reductive acetyl CoA pathway, the 3-hydroxypropionate cycle, or the reverse Krebs cycle, these pathways are relatively small contributors to global carbon fixation compared to that catalyzed by RuBisCO.
The desired reaction is the addition of carbon dioxide to RuBP (carboxylation), a key step in the Calvin–Benson cycle, but approximately 25% of reactions by RuBisCO instead add oxygen to RuBP (oxygenation), creating a product that cannot be used within the Calvin–Benson cycle
The enzyme ribulose-1,5-bisphosphate carboxylase-oxygenase catalyzes the reaction between RuBP and carbon dioxide.The product is the highly unstable six-carbon intermediate known as 3-keto-2-carboxyarabinitol 1,5-bisphosphate, or 2'-carboxy-3-keto-D-arabinitol 1,5-bisphosphate (CKABP). [8]
Calvin–Benson cycle. C 3 carbon fixation is the most common of three metabolic pathways for carbon fixation in photosynthesis, the other two being C 4 and CAM.This process converts carbon dioxide and ribulose bisphosphate (RuBP, a 5-carbon sugar) into two molecules of 3-phosphoglycerate through the following reaction:
Cyanobacteria such as these carry out photosynthesis.Their emergence foreshadowed the evolution of many photosynthetic plants and oxygenated Earth's atmosphere.. Biological carbon fixation, or сarbon assimilation, is the process by which living organisms convert inorganic carbon (particularly carbon dioxide) to organic compounds.
The resulting concentration of carbon dioxide near RuBisCO decreases the proportion of ribulose-1,5-bisphosphate oxygenation and thereby avoids costly photorespiratory reactions. The surrounding shell provides a barrier to carbon dioxide loss, helping to increase its concentration around RuBisCO. [4] [5] [6]
This chemical reaction is catalyzed by the enzyme RuBisCO, and this enzyme-catalyzed reaction creates the primary kinetic isotope effect of photosynthesis. [1] It is also largely responsible for the isotopic compositions of photosynthetic organisms and the heterotrophs that eat them.
C 4 photosynthesis reduces photorespiration by concentrating CO 2 around RuBisCO. To enable RuBisCO to work in a cellular environment where there is a lot of carbon dioxide and very little oxygen, C 4 leaves generally contain two partially isolated compartments called mesophyll cells and bundle-sheath cells.