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Stroma, in botany, refers to the colorless fluid surrounding the grana within the chloroplast. [1] Within the stroma are grana (stacks of thylakoid), the sub-organelles where photosynthesis is started [2] before the chemical changes are completed in the stroma. [3] Photosynthesis occurs in two stages.
Within the envelope membranes, in the region called the stroma, there is a system of interconnecting flattened membrane compartments, called the thylakoids.The thylakoid membrane is quite similar in lipid composition to the inner envelope membrane, containing 78% galactolipids, 15.5% phospholipids and 6.5% sulfolipids in spinach chloroplasts. [3]
The thylakoid membranes of higher plants are composed primarily of phospholipids [5] and galactolipids that are asymmetrically arranged along and across the membranes. [6] Thylakoid membranes are richer in galactolipids rather than phospholipids; also they predominantly consist of hexagonal phase II forming monogalacotosyl diglyceride lipid.
In general outline, photosynthesis is the opposite of cellular respiration: while photosynthesis is a process of reduction of carbon dioxide to carbohydrates, cellular respiration is the oxidation of carbohydrates or other nutrients to carbon dioxide. Nutrients used in cellular respiration include carbohydrates, amino acids and fatty acids.
5 cluster (including two chloride ions), one non heme Fe 2+ and two putative Ca 2+ ions per monomer. [4] There are several crystal structures of photosystem II. [5] The PDB accession codes for this protein are 3WU2, 3BZ1, 3BZ2 (3BZ1 and 3BZ2 are monomeric structures of the Photosystem II dimer), [4] 2AXT, 1S5L, 1W5C, 1ILX, 1FE1, 1IZL.
The electron transfers from pheophytin to plastoquinone (PQ), which takes 2e-(in two steps) from pheophytin, and two H + Ions from the stroma to form PQH 2. This plastoquinol is later oxidized back to PQ, releasing the 2e - to the cytochrome b 6 f complex and the two H + ions into the thylakoid lumen .
The cytochrome b 6 f complex is a dimer, with each monomer composed of eight subunits. [3] These consist of four large subunits: a 32 kDa cytochrome f with a c-type cytochrome, a 25 kDa cytochrome b 6 with a low- and high-potential heme group, a 19 kDa Rieske iron-sulfur protein containing a [2Fe-2S] cluster, and a 17 kDa subunit IV; along with four small subunits (3-4 kDa): PetG, PetL, PetM ...
Light-dependent reactions of photosynthesis at the thylakoid membrane. Photosystems are functional and structural units of protein complexes involved in photosynthesis. Together they carry out the primary photochemistry of photosynthesis: the absorption of light and the transfer of energy and electrons.