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Shelford's law of tolerance is a principle developed by American zoologist Victor Ernest Shelford in 1911. It states that an organism 's success is based on a complex set of conditions and that each organism has a certain minimum, maximum, and optimum environmental factor or combination of factors that determine success. [ 1 ]
A well-known example of a specialist animal is the monophagous koala, which subsists almost entirely on eucalyptus leaves. The raccoon is a generalist, because it has a natural range that includes most of North and Central America, and it is omnivorous, eating berries , insects such as butterflies, eggs, and various small animals.
All species have limits of tolerance to abiotic factors. Too much or too little of anything can lower their survival and reproductive success and cause reduced fitness. Changes in temperature resulting from global warming, for example, may cause a species to change its geographical distribution northward.
Species distribution, or species dispersion, [1] is the manner in which a biological taxon is spatially arranged. [2] The geographic limits of a particular taxon's distribution is its range , often represented as shaded areas on a map.
Bergmann's rule - Penguins on the Earth (mass m, height h) [1] Bergmann's rule is an ecogeographical rule that states that, within a broadly distributed taxonomic clade, populations and species of larger size are found in colder environments, while populations and species of smaller size are found in warmer regions.
Liebig's law has been extended to biological populations (and is commonly used in ecosystem modelling).For example, the growth of an organism such as a plant may be dependent on a number of different factors, such as sunlight or mineral nutrients (e.g., nitrate or phosphate).
The limiting factor also causes competition between individuals of a species population. For example, space is a limiting factor. Many predators and prey need a certain amount of space for survival: food, water, and other biological needs. If the population of a species is too high, they start competing for those needs.
As a result, larger species are able to survive on a lower quality diet than smaller species. For example, grazing animals make up for their poor quality diet by digesting food longer and are able to extract more energy from it (Maurer et al. 1992 [7]). Smaller species tend to specialize in a habitat that can provide them with a high quality diet.