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Unlike xylem (which is composed primarily of dead cells), the phloem is composed of still-living cells that transport sap. The sap is a water-based solution, but rich in sugars made by photosynthesis. These sugars are transported to non-photosynthetic parts of the plant, such as the roots, or into storage structures, such as tubers or bulbs. [14]
Between the xylem and phloem is a meristem called the vascular cambium. This tissue divides off cells that will become additional xylem and phloem. This growth increases the girth of the plant, rather than its length. As long as the vascular cambium continues to produce new cells, the plant will continue to grow more stout.
Sieve cells are long, conducting cells in the phloem that do not form sieve tubes. The major difference between sieve cells and sieve tube members is the lack of sieve plates in sieve cells. [1] They have a very narrow diameter and tend to be longer in length than sieve tube elements as they are generally associated with albuminous cells. [4]
Xylem cells are dead, hard-walled hollow cells arranged to form files of tubes that function in water transport. A tracheid cell wall usually contains the polymer lignin. [citation needed] The phloem, on the other hand, consists of living cells called sieve-tube members. Between the sieve-tube members are sieve plates, which have pores to allow ...
The cambium present between primary xylem and primary phloem is called the intrafascicular cambium (within vascular bundles). During secondary growth, cells of medullary rays, in a line (as seen in section; in three dimensions, it is a sheet) between neighbouring vascular bundles, become meristematic and form new interfascicular cambium ...
The next layer is the cambium, a very thin layer of undifferentiated cells that divide to replenish the phloem cells on the outside and the xylem cells to the inside. The cambium contains the growth meristem of the trunk. [3] Directly inside of the cambium is the sapwood, or the live xylem cells. These cells transport the water through the tree.
According to the hypothesis, the high concentration of organic substances, particularly sugar, inside the phloem at a source such as a leaf creates a diffusion gradient (osmotic gradient) that draws water into the cells from the adjacent xylem. This creates turgor pressure, also called hydrostatic pressure, in the phloem. The hypothesis states ...
The siphonostele shown on the left may also be called an amphiphloic siphonostele. The eustele shown on the right is collateral, i.e. with all the phloem on one side of the xylem. Siphonosteles have a central region of ground tissue called the pith, with the vascular strand comprising a hollow cylinder surrounding the pith. [9]