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Voles are seldom seen outside these runways, which enable a faster and safer locomotion and easier orientation. The climbing ability of the common vole is very poor. Underground nests are dug 30–40 cm (12–16 in) deep into the ground and are used for food storage, offspring raising, and as a place for rest and sleep.
Townsend's vole lives in a burrow system and creates runways among the vegetation in its habitat. The runways are used all year round by successive generations of voles and may be 2.5 to 5 cm (1 to 2 in) deep. [5] In the summer the voles may take advantage of the denser cover available and also move about elsewhere.
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Every year, an individual female beach vole typically produces two litters of three to five offspring, but most will live for less than one year. [6] The beach voles have a habit of building runways above or in the ground under the beach grass, in order to better stay hidden from aerial predators. These runways may contain cut grass.
Eastern meadow voles are active year-round [8] [9] and day or night, with no clear 24-hour rhythm in many areas. [10] Most changes in activity are imposed by season, habitat, cover, temperature, and other factors. Eastern meadow voles have to eat frequently, and their active periods (every two to three hours) are associated with food digestion.
The long-tailed vole are apprehensive of other voles. [12] They are found in areas inhabited by other microtines, but generally avoid contact. [13] The montane vole is a more aggressive animal and is known to displace them from their habitat. [4] The more long-tailed voles in a given area, the more aggressive the montane voles become. [4]