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Phloem (/ ˈ f l oʊ. əm /, FLOH-əm) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as photosynthates, in particular the sugar sucrose, [1] to the rest of the plant. This transport process is called translocation. [2]
In the stems of some Asterales dicots, there may be phloem located inwardly from the xylem as well. Between the xylem and phloem is a meristem called the vascular cambium. This tissue divides off cells that will become additional xylem and phloem. This growth increases the girth of the plant, rather than its length.
The phloem is the living portion of the vascular system of a plant, and serves to move sugars and photosynthate from source cells to sink cells. Phloem tissue is made of sieve elements and companion cells, and is surrounded by parenchyma cells. The sieve element cells work as the main player in transport of phloem sap.
There is also a tissue between xylem and phloem, which is the cambium. The xylem typically lies towards the axis ( adaxial ) with phloem positioned away from the axis ( abaxial ). In a stem or root this means that the xylem is closer to the centre of the stem or root while the phloem is closer to the exterior.
Xylem and Phloem. A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, engages in photosynthesis, stores nutrients, and produces new living tissue. [1]
Sieve elements are specialized cells that are important for the function of phloem, which is a highly organized tissue that transports organic compounds made during photosynthesis. Sieve elements are the major conducting cells in phloem. Conducting cells aid in transport of molecules especially for long-distance signaling.
The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.
The phloem sugar is consumed by cellular respiration or converted into starch, which is insoluble and exerts no osmotic effect. With much of the sucrose having been removed, the water exits the phloem by osmosis or is drawn by transpiration into nearby xylem vessels, lowering the turgor pressure within the phloem. [ 4 ]