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The combination of one or two Clearflight Pied alleles with two Recessive Pied alleles produces the Black-eyed Clear variety, with the appearance described above. This combination appears to result in the complete suppression of the melanin pigment in all the feathers, yet leaves the eyes jet-black.
The eye of the German Fallow is a deep ruby-red, like an Ino's but a shade darker, with the usual white iris ring when adult, [3] [4] and the Scottish Fallow has a pink iris ring. On hatching, young English Fallows have red eyes like Inos but young German Fallow chicks have plum-coloured eyes rather like Cinnamon chicks.
That is, the presence of a single wild-type allele is sufficient to permit the full production and normal distribution of the black melanin pigment. Among the budgerigar fancy such a bird is said to be a Light Green split Scottish fallow, usually written Light Green/Scottish fallow.
The most obvious distinction from Cinnamons is the red eye, which in the German Fallow is a deep ruby-red, like an Ino's but a shade darker, with the usual white iris ring when adult. [2] [3] The English Fallow has an eye of a clear bright red, without a white iris ring—a beautiful and attractive feature. [2]
The stroma around the iris sphincter muscle becomes darker than the peripheral stroma. [medical citation needed] A stimulation of melanin synthesis within iris melanocytes has been postulated. [21] Neoplasm – Nevi and melanomatous tumors. Iridocorneal endothelium syndrome [3] Iris ectropion syndrome [3] [22]
In humans, the pigmentation of the iris varies from light brown to black, depending on the concentration of melanin in the iris pigment epithelium (located on the back of the iris), the melanin content within the iris stroma (located at the front of the iris), and the cellular density of the stroma. [4]
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All pied budgerigars are characterised by having irregular patches of completely clear feathers appearing anywhere in the body, head or wings. These clear feathers are pure white in blue-series birds and yellow in birds of the green series. Such patches are completely devoid of black melanin pigment. The remainder of the body is coloured normally.
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